One day, Usagi Tsukino, clumsy 2nd-year middle school student, stumbles upon a talking cat named Luna. Luna tells her that she is destined to be Sailor Moon, "champion of love and justice", and she must search for the fabled Moon Princess. Usagi finds friends that turn out to be destined senshi as well, and together they fight to save the world from the certain doom brought upon by the Dark Kingdom.

Three decades after the great war between the humans and the Zentradi, the U.N. government is developing new technologies to use in their transforming fighter aircraft by running tests on the colony planet Eden. Military test pilots and former childhood friends, loose cannon Isamu Alva Dyson and the Zentradi mixed race Guld Goa Bowman, are selected to each pilot a new aircraft (Shinsei Industries' YF-19 & General Galaxy's YF-21) for Project Super Nova, to choose the newest successor to the VF-11 Thunderbolt variable fighter which is currently still in use by the U.N. Spacy military forces. Their own personal grudges end up disrupting the tests, and begin to wreak havoc on the program.