The Meiji Era was one of great renewal for Japan, where swords and killing were outlawed. However, many survivors from the time of Revolution still live, lurking in the shadows and waiting for a chance to use their killing blades again. Only Kenshin Himura, formerly one of the most brutal of killers, hopes to keep his swordsman's honor and still live in the new era.

Sakuma Ryu, who runs a liquor store alone in downtown Tokyo, ends up living with his eldest son and his wife after a small fire breaks out. His sons are always there to help, but Ryu, who has stubbornly lived alone since her husband's death, feels uncomfortable there, and frequently visits his tea-drinking buddy Kinue to vent his frustrations. However, Kinue's sudden death and a postcard inspire Ryu to set off on a solo trip in her husband's beloved Mini Cooper. Her destination is Matsuyama in Shikoku, where she spent her childhood and where her first love, for whom she harbored a faint crush, still lives. The Mini Cooper speeds smoothly between Gifu and Kyoto. But when Shota and a young couple from the future join Ryu in the Mini Cooper, things start to look ominous...